Koi or more specifically
nishikigoi, are ornamental domesticated varieties of the common
carp (Cyprinus carpio) that are kept for decorative purposes in outdoor koi ponds or water gardens. They are also called Japanese carp. Koi are among the longest-living vertebrates, with some animals living over 200 years. The most popular category of koi is the
Gosanke, which is made up of the
Kohaku, T
aisho Sanshoku, and
Showa Sanshoku varieties.
The carp is a large group of fish originally found in Central Europe and Asia. Various carp species were originally domesticated in East Asia, where they were used as food fish. The ability of carp to survive and adapt to many climates and water conditions allowed the domesticated species to be propagated to many new locations including Japan. Natural color mutations of these carp would have occurred across all populations. Carp were first bred for color mutations in China more than a thousand years ago, where selective breeding of the Prussian carp (Carassius gibelio) led to the development of the goldfish
Carp are known as koi in Japan. Of the various domesticated carp species, the common carp (Cyprinus carpio) is one of the more commonly used in aquaculture. The common carp was aquacultured as a food fish as at least as far back as the 5th century in China. The common carp was also known to have been aquacultured in Europe by the Roman Empire, which could have spanned a time period of 27 BC to 400 AD. Common carp were first introduced into Japan by way of China between 400 to 600 years ago. Common carp were first bred for color in Japan in the 1820s, initially in the town of Ojiya in the Niigata prefecture on the north eastern coast of Honshu island. By the 20th century, a number of color patterns had been established, most notably the red-and-white Kohaku. The outside world was not aware of the development of color variations in koi until 1914, when the Niigata koi were exhibited in the annual exposition in Tokyo. At that point, interest in koi exploded throughout Japan. The hobby of keeping koi eventually spread worldwide. Koi are now commonly sold in most pet stores, with higher-quality fish available from specialist dealers.
Extensive hybridization between different populations has muddled the historical zoogeography of the common carp. However, scientific consensus is that there are at least two subspecies of the common carp, one from Western Eurasia (Cyprinus carpio carpio) and another from East Asia (Cyprinus carpio haematopterus). One recent study on the mitochondrial DNA of various common carp indicate that koi are of the East Asian subspecies. However another recent study on the mitochondrial DNA of koi have found that koi are descended from multiple lineages of common carp from both Western Eurasian and East Asian varieties. This could be the result of koi being bred from a mix of East Asian and Western Eurasian carp varieties, or koi being bred exclusively from East Asian varieties and being subsequently hybridized with Western Eurasian varieties (the butterfly koi is one known product of such a cross). Which is true has not been resolved.
